May I add that Avery -- a microbiologist and biochemist -- was very reluctant to comment on genetics, especially when his
main consultant was Dobzhansky -- the person who best fits McCarty's attribution.
Had he talked to Muller, Avery would have had much more encouragement! Few geneticists had the training to do much with the
chemical ideas; a few dabbled in cytochemistry or had the eclecticism of an EB Wilson.
It is a pleasure to hear from an economist in this field. Whence your interest?